Evo Vuce i ovde ti se spominje sto ti je Kyros pisao ,Oni spominju 1,36 po generaciji.
"This rise in frequency, if spread evenly over ~34 generations, would require an average increase by a factor of ~1.36 per generation"
http://blogs.discovermagazine.com/g...ect-descendants-of-genghis-khan/#.V-1xk4iLSUk
Gene Expression
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1 in 200 men direct descendants of Genghis Khan
By Razib Khan | August 5, 2010 12:38 am
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473px-YuanEmperorAlbumGenghisPortraitIn 2003 a groundbreaking historical genetics paper reported results which indicated that a substantial proportion of men in the world are direct line descendants of Genghis Khan. By direct line, I mean that they carry Y chromosomes which seem to have come down from an individual who lived approximately 1,000 years ago. As Y chromosomes are only passed from father to son, that would mean that the Y is a record of one’s patrilineage. Genghis Khan died ~750 years ago, so assuming 25 years per generation, you get about 30 men between the present and that period. In more quantitative terms, ~10% of the men who reside within the borders of the Mongol Empire as it was at the death of Genghis Khan may carry his Y chromosome, and so ~0.5% of men in the world, about 16 million individuals alive today, do so. Since 2003 there have been other cases of “super-Y” lineages. For example the Manchu lineage and the Uí Néill lineage. The existence of these Y chromosomal lineages, which have burst upon the genetic landscape like explosive stars sweeping aside all other variation before them, indicates a periodic it “winner-take-all” dynamic in human genetics more reminiscent of hyper-polygynous mammals such as elephant seals. As we do not exhibit the sexual dimorphism which is the norm in such organisms, it goes to show the plasticity of outcome due to the flexibility of human cultural forms.
ResearchBlogging.orgJason Goldman of Thoughtful Animal reminded me of the 2003 paper a few days ago, so I thought it would be useful to review it again for new readers (as I know most of you have not been reading for 7 years!). To understand how one Y chromosomal lineage can have such a wide distribution across such a large proportion of the human race, here is a quote attributed to Genghis Khan:
The greatest joy for a man is to defeat his enemies, to drive them before him, to take from them all they possess, to see those they love in tears, to ride their horses, and to hold their wives and daughters in his arms.
You’re probably more familiar with the paraphrase in Conan the Barbarian.
The Genetic Legacy of the Mongols:
We have identified a Y-chromosomal lineage with several unusual features. It was found in 16 populations throughout a large region of Asia, stretching from the Pacific to the Caspian Sea, and was present at high frequency: ∼8% of the men in this region carry it, and it thus makes up ∼0.5% of the world total. The pattern of variation within the lineage suggested that it originated in Mongolia ∼1,000 years ago. Such a rapid spread cannot have occurred by chance; it must have been a result of selection. The lineage is carried by likely male-line descendants of Genghis Khan, and we therefore propose that it has spread by a novel form of social selection resulting from their behavior.
What is social selection? In this context it’s pretty obvious, the Mongol Empire was the personal property of the “Golden Family,” the family of Genghis Khan. More precisely this came to consist of the descendants of Genghis Khan’s four sons by his first and primary wife, Jochi, Chagatai, Ogedei, and Tolui. Like descent from the gods in the mythology of the Classical World, or the House of David in medieval Christian monarchies, a line back to Genghis Khan became a necessary precondition for fitness to be a ruler in the centuries after the rise of the Mongol Empire across much of Asia.
To me the power and fury of the Mongol expansion, the awe and magnetism which Genghis Khan’s bloodline held for Asiatic societies in the wake of their world conquest, is attested to by the fact that descent from Genghis Khan became a mark of prestige even within Islamic societies. Timur claimed a relationship to Chagatai. His descendants in India, the Timurids, retained pride in their Genghiside heritage. In Russia among the Muslim Tatars and in Central Asia among the Uzbeks descent from Genghis Khan was a major calling card for any would-be warlord. This is peculiar in light of the fact that Genghis Khan, and his near descendants, were non-Muslims! Not only were they non-Muslims, but the Mongol assault on West Asian Muslims societies was particularly deleterious; it is generally assumed that Iran and Mesopotamia’s relatively productive irrigation system were wrecked during the Mongol conquests to the point where it took centuries for them to rebound to their previous levels of productivity. More symbolically, it was the Mongols who finally extinguished the Abbasid Caliphate.
In Muslim societies pride of place is given to Sayyids, descendants of Muhammad through his grandsons Hasan and Husain. Naturally this is often fictive, but that matters little. In fact in the Golden Horde, the northwestern region of the Mongol Empire which eventually gave rise to the Tatars who imposed the yoke on the Russians, non-Genghiside warlords produced fictive genealogies claiming descent from Muhammad as a way to negate the lineage advantage of their Genghiside rivals. But it is still shocking that there was even a question as to whether descent from Genghis Khan was more prestigious than descent from the prophet of Islam!
And the power of descent from Genghis Khan, the monopoly of the commanding heights which his male line descendants still felt to be theirs by right of their blood, obtained at the heart of his Empire, Mongolia, down to a very late period. The last of the great steppe polities, the Zunghar Empire, was defeated by the Manchus in part because it was led by a subset of the Oirat Mongols, a tribe whose leadership were not descended in the male line from the Golden Family, and so could not convince the Genghiside nobility of eastern Mongolia to align with them. From the perspective of moderns, who tend to conceive of historical patterns and forces in economic, or at least ideological, terms, this fixation on blood descent seems ridiculous. I suspect that many pre-modern people, who were accustomed to small family groups and kin networks in a way we are not, would find our own surprise rather perplexing.
So what did they find in the paper? First, they discovered that there was a particular Y chromosomal haplotype, a set of unique genetic markers, which was found across much of Asia. This haplotype seems to have expanded relatively recently, as was evident from small number of mutational steps connecting all of the local variants. Figure 1 illustrates the phylogenetic network:
stardd1
The shaded area represents the star-phylogeny. It’s characterized by a core haplotype, a nearby set of variants separated by one mutational step. This suggests that the genetic variant has risen rapidly in frequency before mutations had time to build up variation and generate a more complex topology. Observe the greater complexity of the network for other Y lineages. Here is the text which explains the factors behind the rise of the Genghis Khan haplotype:
This rise in frequency, if spread evenly over ~34 generations, would require an average increase by a factor of ~1.36 per generation and is thus comparable to the most extreme selective events observed in natural populations, such as the spread of melanic moths in 19th-century England in response to industrial pollution…We evaluated whether it could have occurred by chance. If the population growth rate is known, it is possible to test whether the observed frequency of a lineage is consistent with its level of variation, assuming neutrality…Using this method, we estimated the chance of finding the low degree of variation observed in the star cluster, with a current frequency of 8%, under neutral conditions. Even with the demographic model most likely to lead to rapid increase of the lineage, double exponential growth, the probability was <10−237; if the mutation rate were 10 times lower, the probability would still be <10−10. Thus, chance can be excluded: selection must have acted on this haplotype.
Pogledajte prilog 405109
Could biological selection be responsible Although this possibility cannot be entirely ruled out, the small number of genes on the Y chromosome and their specialized functions provide few opportunities for selection…It is therefore necessary to look for alternative explanations. Increased reproductive fitness, transmitted socially from generation to generation, of males carrying the same Y chromosome would lead to the increase in frequency of their Y lineage, and this effect would be enhanced by the elimination of unrelated males….
A factor of 1.36 per generation is crazy high. In theory of course drift could do this, but in theory the molecules of gas in a room could all congeal to one corner. As noted in the text the Y chromosome is not rich in biologically useful genes. It may be that in the near future we’ll find something peculiar about the carriers of this particular haplotype, but until then, this map speaks for itself:
star3
Citation: ZERJAL, T. (2003). The Genetic Legacy of the Mongols The American Journal of Human Genetics, 72 (3), 717-721 DOI: 10.1086/367774
CATEGORIZED UNDER: GENETICS, HISTORY
MORE ABOUT: GENGHIS KHAN, GENGHIS KHAN GENETICS, MONGOL GENETICS, MONGOLS